Annual or perennial branching herbs, with opposite, usually toothed leaves and rather small, long-peduncled discoid and radiate heads, terminal, or in the upper axils, or rays wanting in some species. Involucre campanulate, its bracts in about 2 series, herbaceous, loosely appressed. Receptacle convex or elongated, chaffy, its chaff embracing the disk-achenes and at length falling away with them. Ray-flowers yellow, or white, pistillate, sometimes wanting. Disk-flowers yellow, perfect, their corollas tubular with an expanded 4-5-cleft limb. Anthers truncate at the base. Style-branches of the disk-flowers long, sometimes penicillate at the summit. Ray-achenes 3-sided, or compressed, those of the disk-flowers compressed, margined. Pappus of 1-3 awns, or more. [Greek, spot- or stain-flower, not significant.]

About 30 species, natives of warm and tropical regions. Type species: Spilanthes urens Jacq.

1. Spilanthes Repens (Walt.) Michx. Spilanthes

Fig. 4441

Anthemis repens Walt. Fl. Car. 211. 1788. Spilanthes repens Michx. Fl. Bor. Am. 2: 131. 1803. S. americana repens A. H. Moore, Proc. Am. Acad. 42: 547. 1907.

Perennial, usually rooting at the lower nodes; stem slender, simple or branched, spreading or ascending, 8'-2° long, pubescent, or nearly glabrous. Leaves ovate to lanceolate, petioled, acute or acuminate at the apex, or the lower obtuse, coarsely toothed, or nearly entire, 1'-3' long; heads long-peduncled, solitary at the end of the stem and branches, 6"-10" broad; bracts of the involucre oblong to oblong-lanceolate, obtuse or acute; rays 8-12, yellow; receptacle narrowly conic; achenes oblong, most of them roughened when mature and hispidulous; pappus of 1 or 2 very short awns, or none.

In moist or wet soil, Missouri to Texas, east to South Carolina and Florida. June-Sept.

1 Spilanthes Repens Walt Michx Spilanthes 1112