Cactus. This word, applied in the form of κάκτος by the ancient Greeks to some prickly plant, was adopted by Linnaeus as the name of a group of curious succulent or fleshy-stemmed plants, most of them prickly and leafless, some of which produce beautiful flowers, and are now so popular in our gardens that the name has become familiar. As applied by Linnaeus, the name Cactus is almost conterminous with what is now regarded as the natural order Cactaceae, which embraces several modern genera. It is one of the few Linnaean generic terms which have been entirely set aside by the names adopted for the modern divisions of the group.

Fig. 1.  Prickly Pear.

Fig. 1. - Prickly Pear (Opuntia vulgaris). 1, Flower reduced; 2, Same in vertical section; 3, Flattened branch much reduced; 4, Horizontal plan of arrangement of flower.

The Cacti may be described in general terms as plants having a woody axis, overlaid with thick masses of cellular tissue forming the fleshy stems. These are extremely various in character and form, being globose, cylindrical, columnar or flattened into leafy expansions or thick joint-like divisions, the surface being either ribbed like a melon, or developed into nipple-like protuberances, or variously angular, but in the greater number of the species furnished copiously with tufts of horny spines, some of which are exceedingly keen and powerful. These tufts show the position of buds, of which, however, comparatively few are developed. The stems are in most cases leafless, using the term in a popular sense; the leaves, if present at all, being generally reduced to minute scales. In one genus, however, Peireskia, the stems are less succulent, and the leaves, though rather fleshy, are developed in the usual form. The flowers are frequently large and showy, and are generally attractive from their high colouring.

In one group, represented by Cereus, they consist of a tube, more or less elongated, on the outer surface of which, towards the base, are developed small and at first inconspicuous scales, which gradually increase in size upwards, and at length become crowded, numerous and petaloid, forming a funnel-shaped blossom, the beauty of which is much enhanced by the multitude of conspicuous stamens which with the pistil occupy the centre. In another group, represented by Opuntia (fig. 1), the flowers are rotate, that is to say, the long tube is replaced by a very short one. At the base of the tube, in both groups, the ovary becomes developed into a fleshy (often edible) fruit, that produced by the Opuntia being known as the prickly pear or Indian fig.

The principal modern genera are grouped by the differences in the flower-tube just explained. Those with long-tubed flowers comprise the genera Melocactus, Mammillaria, Echinocactus, Cereus, Pilocereus, Echinopsis, Phyllocactus, Epiphyllum, etc.; while those with short-tubed flowers are Rhipsalis, Opuntia, Peireskia, and one or two of minor importance. Cactaceae belong almost entirely to the New World; but some of the Opuntias have been so long distributed over certain parts of Europe, especially on the shores of the Mediterranean and the volcanic soil of Italy, that they appear in some places to have taken possession of the soil, and to be distinguished with difficulty from the aboriginal vegetation. The habitats which they affect are the hot, dry regions of tropical America, the aridity of which they are enabled to withstand in consequence of the thickness of their skin and the paucity of evaporating pores or stomata with which they are furnished, - these conditions not permitting the moisture they contain to be carried off too rapidly; the thick fleshy stems and branches contain a store of water. The succulent fruits are not only edible but agreeable, and in fevers are freely administered as a cooling drink.

The Spanish Americans plant the Opuntias around their houses, where they serve as impenetrable fences.

Melocactus, the genus of melon-thistle or Turk's-cap cactuses, contains, according to a recent estimate, about 90 species, which inhabit chiefly the West Indies, Mexico and Brazil, a few extending into New Granada. The typical species, M. communis, forms a succulent mass of roundish or ovate form, from 1 ft. to 2 ft. high, the surface divided into numerous furrows like the ribs of a melon, with projecting angles, which are set with a regular series of stellated spines - each bundle consisting of about five larger spines, accompanied by smaller but sharp bristles - and the tip of the plant being surmounted by a cylindrical crown 3 to 5 in. high, composed of reddish-brown, needle-like bristles, closely packed with cottony wool. At the summit of this crown the small rosy-pink flowers are produced, half protruding from the mass of wool, and these are succeeded by small red berries. These strange plants usually grow in rocky places with little or no earth to support them; and it is said that in times of drought the cattle resort to them to allay their thirst, first ripping them up with their horns and tearing off the outer skin, and then devouring the moist succulent parts.

The fruit, which has an agreeably acid flavour, is frequently eaten in the West Indies. The Melocacti are distinguished by the distinct cephalium or crown which bears the flowers.